page 235

ON SOME MESOZOIC FOSSILS FROM CENTRAL AUSTRALIA.


BY THE REV. J. E. TENISON-WOODS, F.G.S., &C.

WITH TWO PLATES.

 

The fossils described and figured in this paper were obtained by Mr. Gilliatt, one of the surveyors in the Mining Department. They were found in sinking a well on Mount Stewart Run, which is situated on the Grey Ranges. The fossiliferous bed was struck at a depth of 90 feet. The remains comprise many specimens of Avicula, some Grypheoe, a Trigonia, Belemnite, Pecten, Ostrea, Pinna, Cyprina, Mytilus, and many broken fragments. The only species which can be determined, are of Avicula, Pecten, Trigonia and Belemnites, the rest being too imperfect.

The sand is of a greyish green color, with numerous blackish grains. In places it is finely levigated, so as to preserve impressions of the faintest markings on the shells. Some portions are a coarse grey green sand, easily powdered. Under the microscope, this sand is seen to be composed of angular fragments of crystals, with small portions of shelly matter. There were very few rounded grains and none polished. When treated with hydrochloric acid, there was only a slight effervescence, confined I suppose to the fragments of shelly matter. I could not detect any Foraminifera. Some of the fragments of crystals were dark black or green, some opaque white, and some transparent. There were also a few brown opaque grains. The transparent grains seen in polarized light, all showed bands of color and some twin crystals and triclinic felspar. The large green crystals were dark and opaque; some of the smaller dark grains had almost a metallic lustre. From the absence of any glassy particles, as well as there being so little marks of ferric oxides, we may conclude that this sand was derived from the weathering of some of the ancient granite rocks, and not from volcanic or trap rocks. There is a general external resemblance in the rock to the greensands of Britain and France, but the resemblance is merely external. There is no trace of the glanconitic

page 236

materials, to which the color of so many of the European green sands is due. It is owing in that case, to a peculiar green deposit in the chambers and cells of Foraminifera, while in the Australian rocks, there are no Foraminifera and very little lime. The green color is due to small fragments of a material which I believe to be opaque hornblende. There is some admixture of iron pyrites, and a good deal of brown coal and fragments of coniferous wood mixed with fossil remains, but there are no notes to show whether they occurred in distinct bands or were indiscriminately mingled with the rest.

The fossils contained in this collection, comprise: One fine specimen of the guard of a Belemnite, which I regard as identical with Belemnites australis. Moore.* Some valves too imperfect for satisfactory identification, but not unlike Cuculloea inflata. Moore (loc. cit. p. 250). Also fragments of a large Cyprina. (C. expansa Etheridge? Jour. Geol. Soc. 1872, p. 338), a Mya, Tellina, and finally numerous large and well-preserved specimens of Avicula with characters which belong to many of the species described by Moore, besides a single valve of a species of Trigonia which is certainly undescribed. I shall proceed to consider these fossils in detail.

Belemnites australis, Moore (loc. cit. Plate XVI. figs. 1, 2, 3, 4, 5.) Guard hastate, with a rather long, very slightly undulating outline, ventral face flattened but without a trace of a ventral groove; two lateral grooves sharply cut and approximating to the ventral face in the alveolar region, thence bending towards the dorsal aspect with a scarcely perceptible curve and continued in a fine stria on the ventral margin. The specimen is broken round the alveolar cavity, but the extreme length of what remains is 145 minim. width at the alveolar end 20, greatest width at the end of the lateral groove, and about the centre of the fossil 22, ventro-dorsal width greatest at the broken end, and gradually tapering thence to the point.

   * Quart. Jour. Geol. Soc. Lon. 1870, p. 261.

page 237

In the foregoing diagnosis it will be seen that our specimen differs from B. australis in its greater length, in tapering to a point, and the slightly different curvature of the lateral groove. It resembles it in all other particulars, especially in the depression by lateral expansion of the post alveolar region, in the lateral grooves being so sharply cut in the alveolar and post-alveolar region and in the absence of the ventral groove.

I do not therefore think it advisable to erect it into a new species, but should it prove new I propose for it the name of B. oxys. (Pl. XIII., figs. 1, 2 and 3.)

This species belongs evidently to the division Hastati of M. d'Orbigny as Mr. Moore has pointed out in the case of the specimen from Wollumbilla. In this group the guard is elongated and provided with lateral grooves for a portion of the length, but always with a long ventral groove, which is wanting in the Aus­tralian species. The typical species of the group is B. hastatus, Blainville, which is found in all the middle Oolitic beds from the Kellaways rock to Upper Oxfordian. Other members of the same group have been found in the Oolites of France, England, Russia, and the province of Cutch in India. Other forms of the Hastati are found in the Lower Cretaceous, so that the group is consi­dered to belong to the middle and upper mesozoic rocks.

   NEW SPECIES.

Trigonia mesembria, sp. nov. Pl. xii., figs. 1, 2 and 3. T. t. oblongo-ovali, postice producta, tumida, crassa, umbonibus paullum ante-rioribus, prominulis, haud curvatis; margine superiore concavo, elongato, margine posteriore curvato, sed postice truncato; Antice tumide producta, rotundata; sulco posteriore lato, conspicuo, minime profundo, marginibus valde divergentibus a natibus ad marginem; costis transversalibus, haud prominulis, costulis aliquando intercalantibus antice parum undulosis sulco posteriore evanescentibus; carina loevi, sulcis tribus latiusculis munita, lnxula longa loevi ovata, 2 sulcis longis latis, sculpta, basin versus striata; dentibus cardinalibus crassis conspicuis marginibus superioribus bi-sulcatis, sulcis luteralibus 13, crebris, concinnis.

page 238

Shell ovately oblong, produced posteriorly, thick, tumid, very convex, umbones somewhat anterior, not recurved, superior border moderately concave, rather elongate, posterior border curved, slightly truncate at the siphonal margin; anterior border tumidly produced and rounded, posterior groove wide, conspicuous, shallow, widening rapidly from the umbones to the margin, costa irregular, not prominent, some smaller ones occasionally intercalated, all passing horizontally across the valve, slightly undulating anteriorly, disappearing on the posterior groove; siphonal ridge smooth, with three narrow rounded ridges, three rather wide, shallow, grooves between, which become faint as they widen out towards the margin, escutcheon long, ovate, with a flat groove on each side, striae on the lower part, the rest smooth, margins raised; hinge teeth thick, large, prominent, with two grooves on the upper edge and about 13 close, neat, parallel, lateral grooves. The shell is of considerable thickness and is separated into two layers the outer of which is the thicker.

This fossil manifestly belongs to that section of the large genus Trigonia, which is classed as Glabroe or smooth. The section is characterised by sub-quadrangular or elongately ovate shells moderately compressed, area not margined or indicated by a distinct groove, sides ornamented with smooth concentric ribs, sometimes extending over the area, or becoming quite obsolete before reaching it. The type is T. longa, Agassiz.

The group of Glabroe is decidedly Cretaceous, most of the species belong to that period, though one or two species rarely extend into the Jura.

The above fossil comes nearest in its form to T. excentrica, Park, and T. duncombensis, Lycett, both Greensand species or Lower Cretaceous in Britain. There is a single elongated species of the Glabroe division, from the Cretaceous rocks of America. It comes from Columbia. Trigonia semiculta, Stol., of the Cretaceous rocks of Pondicherry, is a short globose species, with the trans­verse costa interrupted about the middle of the shell by the usual smooth anti-carinal space, but it is not grooved as in the present species, is short, has not the posterior carina, and has the costa much more regular.

page 239

It may be as well here to review all the described Australian mesozoic species of Trigonia. They are T. costata, Clarke, which is the same as T. morei, Lycett, T. lineata, Moore, T. nasuta, Etheridge.

The name T. costata, was applied by the Rev. W. B. Clarke, F.R.S., to a species from Western Australia (Greenough River), under the belief that it was Trigonia costata of Parkinson.* In Moore's paper already referred to, the differences between the species are pointed out by the late Mr. Lycett, and the Australian one, named T. moorei. It is one of the group called Costatoe, and therefore entirely different from the new fossil described here.

Trigonia lineata, Moore (loc. cit. p. 255) was described from two imperfect specimens, one a cast showing the teeth, and the other much abraded. It is a gibbous form, as broad as long, and there­fore quite distinct from our species. Professor McCoy placed it with the Permian and Rhaetic genus Myophoria, but Mr. Moore considered that the absence of the oblique keel and the acute posterior side, precluded such a reference.

Trigonia nasuta, Etheridge (Jour. Geol. Soc. Lond. 1872, p. 339, pl. xix., figs. 2, 2a). This fossil was described from a cast only, but which Mr. Etheridge says, belongs to a type unknown in Jurassic rocks. It is much higher and deeper than our fossil, and was referred to the group Scabroe nearest to T. sanctoe crucis, Pictet and Camp., but a discovery of the external surface, might show it to be much nearer to the present species.

Pecten psila, sp. nov. P. t. parva, trigonali-orbiculari, oequivalva, valde depressa, vix convexiuscula, omnino loevi, vel marginem versus tenuissime concentrice striata, auribus inoequalibus, radiatim costatis, antice elongata triangulala producta, posteriore flabelliformi.

Shell small, suborbicular, equivalve, depressed, hardly convex, entirely smooth, and without ornament of any kind, except one or two very faint lines of growth near the exterior margin; ears

   * Organic Remains. vol. 3, pl. 12, fig. 4; Paleontograph Soc. Monograph, Lycett and Moore, Mollusca of the Great Oolite, pl. 5, fig. 23; Lycett. Brit. Foss. Trigonia, vol. 33, p. 227, a wood cut of T. Moorei.

page 240

unequal, radiately ribbed, the anterior one in the form of an elongated triangle, produced along the dorsal margin for half the length of the shell, posterior smaller, fan-shaped.

This small, smooth, depressed Perten, belongs to a type which is common in the mesozoic rocks, the middle mesozoic especially. In a genus so abundantly represented in species during almost every geological period resemblances can be found on every side. But it is particularly like P. socialis, Moore, which is sub-orbicular shell moderately convex, umbones convex and pointed, auricles unequal, anterior one much the largest. Mr. Moore says that the external ornamentation of the shell is not well preserved, but it appears to have been nearly smooth, without visible concentric striae but with depressed radiating ribs. In general form it is not unlike P. rigidus.* It is one of the most abundant shells in the boulders from Wollumbilla, many specimens of various ages appearing on their fractured surfaces.

The fossil I have described might well be a young state of Mr. Moore's fossil. It is much smaller, the average dimension from hinge to margin is about 6 millimeters with a tranverse diameter of 5. It is found abundantly scattered over some fragments of dark, olive greenish marl, looking not unlike Nummulites. As I have only seen the figures and descriptions of P. socialis, and as the details are very imperfect, I think it better not to make too hasty an identification but point out the resemblance for future enquiry.

Avicula barklyi, Moore, vel reflecta, Moore var. gilliatti, pl. xii., figs. 4, 5 and 6. The great mass of the fossils in this collection is a large species of Avicula of which a figure is given. It possesses all the characters of both the fossils above-named in different specimens and I think that the specific distinctions between the two will be found on examination to disappear. Any one who examines the series figured by Mr. Moore from Wollumbilla including eight species, will he inclined to refer them all to one, differing from each other merely in size and mode of growth. The posterior wing auricle is lengthened and undulating in the large and old specimens

   * A fossil species of the Great Oolite, Wiltshire. See Sowerby's Mineral Conchology, plate 205, fig. 8.

page 241

(those of about 90 millimeters long), and the produced posterior wing makes the transverse and longitudinal diameters quite equal. A. barklyi, has been recognized by Prof. Tate, from Cape Creek, and other fossils described by Moore as associated with that fossil from Primrose Springs and Lake Eyre, in Central Australia. (See Trans. Roy. Soc. S. Australia, Vol. VI., p. 144.) They were scattered on the surface much in the way the fossils are at Wollumbilla.

   SUMMARY.

From the occurrence of some cretaceous forms in this deposit, I am inclined to think that it belongs to the so-called great Cretaceous formations of Central Australia, and North-eastern Australia. There is certainly also a mixture of jurassic types, but their evidence is not so marked, and I am inclined to think that there has been a mixture of fossils from different localities in Mr. Moore's collec­tion. There are true Oolitic beds near the Peak Downs, and amongst these I have recognized some of Moore's species, but never the Aviculae, and Belemnite here referred to. It is possible, how­ever, that our Australian Cretaceous rocks may be very low in the series, and contain moreover as in the Australian strata a fusion of those well-marked specific boundaries which paleontologists are accustomed to elsewhere. In any case I am rather disposed to regard the Central Australian beds about Lake Eyre, the Peake and Primrose Springs as cretaceous. I propese to examine the Wollumbilla beds, carefully at my earliest leisure and then will try for its position more certainly.

   EXPLANATION OF PLATES,

   Plate XII.

Fig. 1. - Trigonia mesembria, inner surface of valve.
Fig. 2. - Upper surface showing concentric costa.
Fig. 3. - Anterior end with tooth.
Fig. 4. - Avicula barklyi, upper valve.
Fig. 5. - Ditto ditto, lower valve of same specimen.
Fig. 6. - Another upper valve with better defined ribs.
      All figures reduced one-half nat. size.

   Plate XIII.

Fig. 1. - Belemnites australis, Phillips? or B. oxys, nobis. ventral aspect.
Fig. 2. - Ditto lateral aspect showing the slight curve of the lateral groove.
Fig. 3. - End view with alveolar cavity, lying on ventral surface.
      All figures two-thirds natural size.